Heat oven to 350ยบ F. Spray 13x9-inch pan with cooking spray.
In large mixing bowl, beat cake ingredients with electric mixer on medium approximately 2 minutes, scraping the bowl as necessary. Pour into pan.
Bake 28 to 30 minutes or until toothpick inserted in center of cake comes out clean. Cool cake in pan on cooling rack for 10 minutes.
Filling
With handle of wooden spoon (approximately 1/2" in diameter), poke holes all over cake every 1/2 to 1 inch. Note: Wiping spoon handle with paper towel to reduce sticking may be necessary.
In a medium bowl, pour boiling water over strawberry JELL-O mix; stirring until gelatin is completely dissolved. Stir in cold water. Pour gelatin mixture carefully and slowly over the cake. Refrigerate for 2-3 hours.
Frosting
Beat together lemon pudding mix, milk and lemon zest in a large mixing bowl until blended. Fold in container of cool whip. Spread over cooled cake.
Refrigerate cake for approximately 30 minutes before serving.